The San Diego Zoo's oldest resident, a Galapagos tortoise named Gramma, died Nov. 20. She was estimated to be 141 years old.

After more than a century of munching on her favorite foods of romaine lettuce and cactus fruit, beloved Galapagos tortoise ...

Zoo officials said she came from the Bronx Zoo in either 1928 or 1931 as part of their first group of Galápagos tortoises.

San Diego Zoo staff and animal lovers are reeling this week following the death of beloved Gramma the Galápagos tortoise, by ...

Holiday Lights occupies 12 of the Bronx Zoo’s 265 acres of hardwood forest – an oasis of wilderness amid the Bronx’s urban ...

Gramma the Galápagos tortoise died on Nov. 20, 2025. The oldest animal at the San Diego Zoo was believed to be around 141 ...

London Zoo, previously known as ZSL London Zoo or London Zoological Gardens and sometimes called Regent's Park Zoo, is the ...